First time posting in this forum. I was told this is the best place to ask for advice on sexing plecos.
Would you be able to help me?
These L333s are about 3.5"-4" long.
Here's a top view:

Let's say we go clockwise starting from top (#1), bottom-right (#2), bottom-middle (#3), bottom-left (#4).
Here are some close-ups:
#1

#2

#3

#4

I usually look at the odontodes on the pectoral fins. Going by that, I think, #1 is male, #2 is female, #3 is male, and #4 is female.
I know males occupy caves all the time but all four are usually in caves because I have a bare-bottom tank, several caves and not enough of other cover. Then a friend brought up the body shape and that's when I got confused.
What do you think?
Thanks in advance!!!